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SYNOPSIS
In Piqua, Ohio, George Beard (Kevin Hart) and Harold Hutchins (Thomas Middleditch) are two fourth-grade friends and next-door neighbors who often bring joy to their school, Jerome Horwitz Elementary School, by excessively pranking the cruel teachers, especially the ill-mannered misanthropic principal Benjamin "Benny" Krupp (Ed Helms), which puts them at odds with him. The duo also create comic books about Captain Underpants, a superhero who fights crime while wearing only underwear and a cape. They sell these to their schoolmates through a comic company called Treehouse Comix Inc, housed in their treehouse. George and Harold's pranks come to an apparent end after they tamper with a toilet invention called the Turbo Toilet 2000 made by the school's local snitch, intellectual Melvin Sneedly (Jordan Peele). This causes Krupp to decide to separate the boys and annihilate their friendship.
To prevent this, George hypnotizes Krupp with a 3D Hypno Ring he received out of a cereal box; the boys first order their principal to act like a chicken and a monkey. Finally, they see that Krupp bears a resemblance to Captain Underpants without his toupee and command him to be Captain Underpants. The boys soon learn the severity of their acts when Captain Underpants begins causing some problems around Piqua and take him to their tree-house. There they discover that they can turn Captain Underpants back into Krupp by splashing water on him and can turn him back by snapping their fingers. Believing that Krupp will continue with his plan to separate them, they decide to settle with Captain Underpants but insist that he be dressed up as Krupp under the pretense of a secret identity to which Captain Underpants agrees. His sudden personality change even manages to attract the attention and affection of the school's shy lunch lady Edith (Kristen Schaal).
Just when George and Harold believe that their troubles have ended, Jerome Horwitz Elementary School is visited by an odd, German-accented scientist named Professor Pee-Pee Diarrheastein Poopypants, Esq. (Nick Kroll). Captain Underpants (disguised as Krupp) hires him to be a new teacher, but George and Harold are suspicious of him. As it turns out, Poopypants is seeking to get rid of laughter altogether due to the fact that people have made fun of his name for years. He recruits Melvin to help him as his brain is incapable of having fun.
Soon, Professor Poopypants tries to take over the town with a giant version of the Turbo Toilet 2000, fueled by the school cafeteria's rotten leftovers left out by Edith, and uses Melvin's brain to turn the children into glum, humorless zombies. Captain Underpants tries to stop the villain, but due to having no actual superpowers, is effortlessly defeated and thrown into the toilet. George and Harold are captured and turned into zombies, but are able to break the trance and escape when their laughter damages the Turbo Toilet 2000's computer. Upon consuming the mutated leftovers, Captain Underpants acquires superpowers and with George and Harold's help defeats and shrinks Poopypants, though he escapes shortly thereafter.
Knowing that they cannot control Captain Underpants, George and Harold destroy the Hypno Ring in an attempt to permanently change him back into Krupp. Feeling that Krupp would be nicer if he had friends, the boys set him and lovesick Edith up on a date, thus making Krupp have a change of heart; he returns the comics he took away from George and Harold, and even admits their comics are funny. However, the toxic waste from the Turbo Toilet 2000 transforms all the toilets into vicious monsters that attack the restaurant at which Mr. Krupp and Edith are dining. Upon snapping his fingers (accidentally), Mr. Krupp once again becomes Captain Underpants, carrying George and Harold away to help him fight them, much to Edith's surprise and admiration.
In a mid-credits scene, George and Harold realize that the secretary that they put on the phone before by faking a $1 billion contest has been on the phone for the entire movie. The secretary accidentally hangs up and gets angry, while George and Harold make a new comic based on her reaction.
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